The Babylonian province of Phoenicia and its neighbors passed to Achaemenid rule with the conquest of Babylon by Cyrus in 539/8 BC. The Syro-Phoenician coastal cities remained under Persian rule for the following two centuries. The Phoenician navy supported Persia during the Greco-Persian War (490-49 BC). But when the Phoenicians were overburdened with heavy tributes imposed by the successors of Darius I, revolts and rebellions resumed in the Lebanese coastal cities. Phoenicia eventually fell to Alexander the Great, king of Macedonia in 4th century BC.